Scott beat me to it, I've had trackage like that, it is due to the wood drying out, in my case due to the furnace running in the winter. It dries the air very much, and like most basements it is quite humid in the summer. The wood contracts as it dries, pulling those track nails closer together. I pulled the nails as required, used a dremel cutoff disc to cut a gap, usually that was enough to shorten the rail sufficiently. Added drops if required. When the wood swells those nails will go back to where they were and you'll have some gaps. I painted my wood subroadbed after that but that doesn't completely prevent the problem. I used regular latex paint, perhaps something specifically for sealing would work better. I found the problem existed primarily where track nails extend into the wood. My handlaid track, spiked into homasote, never has had a problem. Some handlaid track for a track in the street scene, where I spiked directly to some pine, had the same problem. I haven't tried it, but avoiding track nails and glueing instead might help. I now only lay track during the heating months. I haven't experienced any problems with excessive gaps opening, yet. Hope this helps.
Gary